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CARe Graduate School’s scholarships

Our call for proposals is now open!

Each year, the CARe Graduate School offers the possibility to international students who have been selected to join the Master’s “Bio-santé” on Cancer, Ageing and Rejuvenation to benefit from a two-year scholarship.

Objective of the call: Allow international CARe students to travel, fund their accommodation, personal expenses and internships during their Master’s studies at the University of Toulouse 3 – Paul Sabatier.

Amount of the scholarship: 7 000 Euros per academic year (14 000 Euros for two years).

The scholarship will include 4 050 Euros per year for accommodation and personal expenses, and 2 950 Euros per year of part-time internship gratuities in France or abroad (paid separately). A round-trip air ticket will also be covered.

Selection criteria:

Students must:

  • Hold at least a Bachelor’s Degree in biology, mathematics, computer science, chemistry, physics or related field from a foreign university;
  • Have been selected to join the Master’s “Bio-santé” on Cancer, Ageing and Rejuvenation.

A strong preference will be given to candidates living abroad at the moment of their application and travelling to France to register to CARe’s Master.

The second year of the scholarship is subjected to the validation of the first year of the Master’s Degree.

Application process:

After receiving a positive response to their application to the Master’s “Bio-santé” on Cancer, Ageing and Rejuvenation, international students shall send a letter of interest by email to Clémence Grosnit ( and Claire Mendoza (, by June 30, 2022.



External offers


PhD thesis in biomathematics: Morphogenetic emergence across biological scales

Centre for Integrative Biology (CBI), Toulouse, France

Description of the project

During development, dynamic interactions between cells allow the emergence of organ shapes. Because of the diversity of cell properties and behaviors, such as migration, division, or adhesion, complex morphogenetic processes have been difficult to tackle experimentally. We study the development of the posterior part of the vertebrate embryo as it contains progenitor cells that will both specify and form distinct tissues, the neural tube (future nervous system), and the paraxial mesoderm (future vertebrae, muscle, and derm). Using time-lapse movies, we will quantify quail embryonic cells and tissues behavior by image analysis. Based on these quantifications, we will develop agent-based and continuum (PDE) models to understand the relationships between the different levels of organization of our system: cell/cell, cell/tissue, and tissue/tissue interactions. These models will be analyzed theoretically and numerically (qualitative behavior, link between the different models) and tested against our biological data. Altogether, we will be able to combine unique experimental results and theoretical/numerical approaches to understand how distinct cellular behaviors synergize to build embryonic tissues and structures.


Morphogenesis, cell and tissue behavior, self-organization, agent-based modeling, continuum modeling, partial differential equations, numerical simulation, scaling/hydrodynamic limits, free boundary problem, time-lapse imaging, migration.


Romanos M, Allio G, Roussigné M, Combres L, Escalas N, Soula C, Steventon B, Trescases A.,

Bénazéraf B.. Cell-to-cell heterogeneity in Sox2 and Bra expression guides progenitor motility and

destiny. Elife. 2021 Oct 5;10.

Degond P, Hecht, S, Romanos, M and Trescases, A. Multi-species viscous models for tissue growth: incompressible limit and qualitative behaviour

Conditions and application

The candidate will work in a highly interdisciplinary collaborative context. Image and data analysis will be performed at the Centre for Integrative Biology (CBI) in Toulouse under the supervision of Bertrand Benazeraf. The theoretical side will be supervised by Ariane Trescases at the Institute of Mathematics of Toulouse (IMT).

Candidates must have a Master’s degree or equivalent in mathematics or a subject with a strong mathematical component (e.g. physics, bio-physics, bio-mathematics, engineering, computer science), skills in programming, and experience and/or demonstrate a strong interest in image analysis and morphogenesis. Funding: CNRS PRIME MITI (three years); Start date: Sept/October 2022.

For application send a unique PDF file containing a CV, mark reports since the beginning of higher education, and a motivation letter with 2 reference contacts to and


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Contact us


CARe Graduate School,
4 bis av. Hubert Curien, 31100 Toulouse, FRANCE


+33 7 64 54 13 99 / +33 7 64 72 12 28